botany term for opposite of pioneer species
I was reading about pioneer species a.k.a. ruderal species, forest dynamics, ecological succession (primary, secondary), climax community, old growth. My question is, what is the best term for the opposite of a pioneer species, a species that always appears later? For example, the Canadian Hemlock. The closest I came so far was "shade tolerant", but I was hoping there might be a term like "straggler species".
Is there a good term for that?