My thought would be lack of humidity from the heated greenhouse, either that or your heated greenhouse was on the fritz one cool night.

Looks like spider mites. Check underside of leaves.

Good luck

Hard to diagnose from here, but I think I would hit them with a pesticide and fungicide to give you a better chance to remedy what is ailing them.

Bill

It also looks like mites to me, check the leaves for very fine webbing and tap a leaf onto a white piece of paper, if minute little specks fall off and move you have mites. The kind of light streaky pattern on that one leaf looks suspicious.

Its good to cover all bases though as Bill suggested and use a good fungicide and pesticide.